Rhachiberothidae

Thorny lacewings Temporal range: Early Cretaceous - Recent
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Neuroptera
Suborder:	Hemerobiiformia
Superfamily:	Mantispoidea
Family:	Rhachiberothidae
Genera
Several, including: Mucroberotha Rhachiberotha and see text

The thorny lacewings, Rhachiberothidae, are a family of winged insects of the order Neuroptera.

Long included in the Berothidae (beaded lacewings), they are usually considered a separate family by current authors. Another closely related group are the mantidflies (Mantispidae), and at least some of the probably paraphyletic group of fossil forms collectively called "Mesithonidae" also seem to be quite close.^[1]

A considerable number of prehistoric genera have been described from fossils dating back to the Early Cretaceous:

Alloberotha
Chimerhachiberotha
Eorhachiberotha
Oisea
Paraberotha - formerly in Mantispidae
Retinoberotha - formerly in Mantispidae
Rhachibermissa
Spinoberotha
Whalfera - formerly in Mantispidae

Footnotes

↑ See references in Haaramo (2008)

References

Haaramo, Mikko (2008): Mikko's Phylogeny Archive: Neuroptera. Version of 2008-MAR-11. Retrieved 2008-APR-27.

Insecta orders

Kingdom: Animalia
Phylum: Arthropoda
(unranked): Pancrustacea
Subphylum: Hexapoda

Extant

Monocondylia	Archaeognatha (jumping bristletails)

D
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Apterygota *	Thysanura (Zygentoma) (silverfish, firebrats)

P
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Palaeoptera

Ephemeropteroidea	Ephemeroptera (mayflies)

Odonatoptera	Odonata (dragonflies, damselflies)

N
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Polyneoptera

Dictyoptera	Blattodea (cockroaches, termites) Mantodea (mantises)

E
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b
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Paraneoptera *	Psocodea (barklice, lice) Thysanoptera (thrips) Hemiptera (cicadas, aphids, true bugs)

E
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basal

Hymenoptera (sawflies, wasps, ants, bees)

Neuropteroidea

Coleopterida	Strepsiptera (twisted-winged parasites) Coleoptera (beetles)

Neuropterida	Raphidioptera (snakeflies) Megaloptera (alderflies, dobsonflies, fishflies) Neuroptera (net-winged insects: lacewings, mantidflies, antlions)

Panorpida
(Mecopterida)

Antliophora	Mecoptera (scorpionflies) + Siphonaptera (fleas) Diptera (gnats, mosquitoes, flies)

Amphiesmenoptera	Trichoptera (caddisflies) Lepidoptera (moths, butterflies)

Four most speciose orders are marked in bold

Italic are paraphyletic groups

Based on Sasaki et al. (2013)

Extinct

Alienoptera Archodonata Blattoptera Caloneurodea Campylopteridae Carbotriplurida Coxoplectoptera Diaphanopterodea Eoblattodea Eocoronidae Eudiaphanoptera Erasipteridae Gerarus Geroptera Glosselytrodea Heraridea Hypoperlida Lapeyriidae Meganisoptera (griffinflies or "giant dragonflies") Megasecoptera Mendozachorista Miomoptera Monura Palaeodictyoptera Paoliida Permoplecoptera Protanisoptera Protelytroptera Protephemerida Protodiptera Protorthoptera Protozygoptera Pseudopulex Rhyniognatha Syntonoptera Titanoptera Triadophlebioptera

Extinct incertae sedis families and genera are marked in italic

Wikispecies

Extant Neuroptera families

Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Subclass: Pterygota
Infraclass: Neoptera
Superorder: Endopterygota

Basal

Osmylidae (osmylids) Nevrorthidae

Suborder Hemerobiiformia

Ithonioidea	Ithonidae (moth lacewings)

Chrysopoidea	Chrysopidae (green lacewings)

Hemerobioidea	Hemerobiidae (brown lacewings)

Coniopterygoidea	Coniopterygidae (dustywings) Sisyridae (spongeflies or spongillaflies)

Mantispoidea	Dilaridae (pleasing lacewings) Mantispidae (mantidflies) Rhachiberothidae (thorny lacewings) Berothidae (beaded lacewings)

Suborder Myrmeleontiformia

Nemopteroidea	Psychopsidae (silky lacewings) Nemopteridae (spoonwings)

Myrmeleontoidea	Nymphidae (split-footed lacewings) Myrmeleontidae (antlions) Ascalaphidae (owlflies)

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